I'd call Winchester. I've always used 2 piece bases (Talley Lightweight) and there isn't any difference. But my references show long action, short action, and WSM for one piece bases and rails. Maybe Winchester has changed something.
The short action and WSM stocks are interchangeable, or at least used to be. I had a 300 WSM for a while and stocks, as well as bottom metal were all the same. As were 2 piece bases, but I've never tried using a rail.